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Why fish size figures disagree

Two sources giving a fish as 22 cm and 40 cm may both be right and may not disagree at all. Four separate distinctions are routinely collapsed in aquarium writing — measurement convention, maximum against typical, wild against captive, and what kind of document the figure came from — and untangling them resolves most apparent conflicts and leaves a small number of real ones.

Short answer

Because they are usually measuring different things. Standard length excludes the tail fin and total length includes it; a maximum is the largest specimen anybody recorded and a typical size is what a well-kept adult reaches; and a taxonomic checklist records the biggest fish its compiler examined, which is not the same as the biggest the species gets.

Standard length and total length

Standard length runs from the snout to the base of the tail fin. Total length runs from the snout to the tip of it. For a fish with a short tail the difference is ten per cent; for one with a long trailing caudal it can be a third of the animal.

Museum and taxonomic sources overwhelmingly publish standard length, because it is reproducible on a preserved specimen whose fins may be damaged. Angling records and aquarium literature overwhelmingly publish total length, because it is what you get when you lay a fish against a ruler.

So a museum figure and a shop figure for the same species will differ, systematically, in the same direction, without either being wrong. AquariumHQ records the measurement type on every size value where the source states one, and records that the source did not state one where it did not — because '40 cm' of an unstated kind is a different fact from '40 cm TL'.

Standard length, fork length and total length on one fishA fish shown in side view with three measurement lines beneath it. Standard length runs from the snout to the base of the tail fin. Fork length runs from the snout to the fork of the tail. Total length runs from the snout to the tip of the longest tail lobe. A note records that for a fish with a long trailing caudal fin the difference between standard and total length can be a third of the animal's stated size.SL — standard lengthFL — forkTL — totalA source that gives a length without saying which of these it used has given an approximation.
Three ways to measure one fish. Museum sources overwhelmingly publish standard length; angling records and aquarium literature publish total length; and the difference can be a third of the animal. Source: AquariumHQ size model. See the guide on why size figures disagree.

Maximum, typical, and the gap between them

A maximum recorded size is one extreme specimen. A typical adult size is what a well-kept fish reaches. The gap is usually large and occasionally decisive.

The goldfish is the clean example. USGS NAS gives a maximum of 410 mm SL and a typical 120–220 mm SL in the same sentence, and separately reports an angling record of 59 cm TL. Those three numbers span a factor of five and every one of them is correct about a different question.

Fishes of Australia publishes both, routinely and explicitly: 'To 7 cm SL, commonly to 4 cm' for the Pacific blue-eye. Almost no aquarium source does this, and its absence is why 'maximum size' has quietly become the number the trade quotes for everything.

Four figures for one fish, all correct
FigureWhat it answersWhere it comes from
41 cm SLThe largest specimen recordedField guide, standard length
12–22 cm SLWhat one usually isSame field guide, same page
59 cm TLThe largest ever caught on a lineAngling record body
20–35 cmWhat a pond fish reaches in practiceAquariumHQ's own position

A checklist is not a survey

This is the distinction that produced the sharpest disagreement AquariumHQ found this sprint. USGS NAS gives the jaguar cichlid as 22 cm SL, crediting a taxonomic checklist. AquariumHQ publishes 30–40 cm. That is roughly a factor of two and it is not a standard-versus-total-length artefact.

The likeliest reconciliation is that a checklist records the largest specimen its compiler examined — a museum collection, assembled for taxonomy rather than for size records — while the aquarium figure describes a heavily farmed fish that has been seen much larger. That is a reasonable story and it has not been established, so both figures are published and the disagreement is left open rather than resolved by picking the convenient one.

Two sources agreeing may be one source twice

USGS NAS gives the jack dempsey as 25 cm SL and credits Kullander (2003). It gives the convict cichlid as 10 cm SL and credits Kullander (2003). It gives the jaguar cichlid as 22 cm SL and credits Kullander (2003). Those figures also stand behind other databases' entries for the same fish.

Two databases agreeing on a number is not two measurements agreeing. It can be one measurement reported twice, and from the outside the two look identical. AquariumHQ records the upstream reference a source credits, wherever it credits one, and refuses to count two values as independent when they name the same work.

What to do with a size figure you have just found

Four questions, in order. They resolve most of it.

  • Is it standard length or total length? If the source does not say, treat the figure as approximate rather than assuming.
  • Is it a maximum or a typical? A maximum is not a planning figure and is quoted as though it were constantly.
  • Is it wild or captive? Farmed stock of several species runs consistently smaller than wild; a few run larger.
  • What kind of document is it? A taxonomic checklist, an angling record and a care sheet are answering three different questions with the same unit.